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Animal Diversity - Invertebrates

The course covers the staggering multitude of invertebrates that make up more than 95% of the animal species.

A thorough survey of the current phylogenetic hypotheses that help making sense of animal diversity is a central theme of the course The course will review data from comparative anatomy, developmental biology, natural history, genetics, genomics, and other fields in order to assess our current understanding of animal evolution. A twelve day field course at a marine lab provides first hand encounters with many of the major animal groups. After the course you will be familiar with the main animal groups, their biology, and current phylogenetic hypotheses detailing their evolution.

  • Course structure

    Teaching format

    Lectures will be on Zoom and material (presentation pdfs) will be available in most cases. In addition we will uses compendia and journal articles.

    There will be a field course held at Tjärnö Marina Laboratorium, Strömstad for about two weeks in November. During the field course you will have the opportunity to participate in sampling trips with R/V Nereus. We will also make excursions to the shore and to freshwater habitats.

    Active participation in group discussions, the field course, submission of a project report and a presentation for the seminar are all compulsory. Teachers can be contacted by email anytime during his/her block of teaching (see schedule for contact information).


    There will be a practical test during the field course. The final examination consists of a formal test with questions which will be distributed by e-mail, and the completion of an individual project.


    Ulf Jondelius

  • Schedule

    The schedule will be available no later than one month before the start of the course. We do not recommend print-outs as changes can occur. At the start of the course, your department will advise where you can find your schedule during the course.
  • Course literature

    Note that the course literature can be changed up to two months before the start of the course.

    Brusca, R.C. et al. Invertebrates. 3:d ed. Sinauer Associates 2016. ISBN 9781605353753. Other materials will be distributed during the course.

  • Course reports

  • Contact

    Student office
    Study counsellor for advanced level