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Animal Diversity - Invertebrates

The course covers the staggering multitude of invertebrates that make up more than 95% of the animal species.

A thorough survey of the current phylogenetic hypotheses that help making sense of animal diversity is a central theme of the course The course will review data from comparative anatomy, developmental biology, natural history, genetics, genomics, and other fields in order to assess our current understanding of animal evolution. A twelve day field course at a marine lab provides first hand encounters with many of the major animal groups. After the course you will be familiar with the main animal groups, their biology, and current phylogenetic hypotheses detailing their evolution.

  • Course structure

    Teaching format

    Lectures will be on Zoom and material (presentation pdfs) will be available in most cases. In addition we will uses compendia and journal articles.

    There will be a field course held at Tjärnö Marina Laboratorium, Strömstad for about two weeks in November.

    Active participation in group discussions, the field course, submission of a project report and a presentation for the seminar are all compulsory. Teachers can be contacted by email anytime during his/her block of teaching (see schedule for contact information).

    Learning outcomes

    A list of the learning outcomes can be found in the syllabus. Please find the link to the syllabus on the right side of this page.


    There will be a practical test during the field course. The final examination consists of a formal test with questions which will be distributed by e-mail, and the completion of an individual project.


    Ulf Jondelius


    Tel: +46 70 7838711

  • Schedule

    The schedule will be available no later than one month before the start of the course. We do not recommend print-outs as changes can occur. At the start of the course, your department will advise where you can find your schedule during the course.
  • Course literature

    Note that the course literature can be changed up to two months before the start of the course.

    Brusca, Moore, Shuster. 2016. Invertebrates 3rd edition. Sinauer Associates.  ISBN 9781605353753.  Other materials will be distributed during the course.

  • Course reports

  • More information

    Field course at the Tjärnö Marina Laboratorium

    During the field course you will have the opportunity to participate in sampling trips with R/V Nereus. We will also make excursions to the shore and to freshwater habitats. It is a good idea to bring warm clothing and rain gear. You will also need to bring bed linen.

    Tjärnö has a comprehensive and compulsory meal plan. The cost for all meals from dinner Nov 15 through breakfast Nov 26 is 2 253  SEK to be paid to Stockholm Unversity bankgiro 5050-0206. Please include the reference number 4946017/460/3322 and your name with the payment. Send me an e-mail at if you have dietary requirements, eg. food allergies, vegetarian diet etc. There is no charge for accommodation at
    Tjärnö. Please arrive at Tjärnö in time for dinner at 17.00 on the first day.

    You will need to book and purchase your ticket individually. Ticket costs for the return trip from Stockholm will be reimbursed by the Stockholm University Department of Biology Education upon presentation of receipts/tickets (contact

    Suggested itinerary (2021) if you are travelling from Stockholm on public transportation:
    15 Nov
    Train 425 from Stockholm Central station to Göteborg 08.30-11.35
    Train 3750 from Göteborg Central Station to Strömstad 12.40-15.30
     Bus 891 from Strömstad station to Tjärnö Lovéncentret 16.05-16.28
    26 Nov
    Bus 891 from Tjärnö Lovéncentret to Strömstad station 07.38-08.05
    Bus 871 from Strömstad to Torp 09.20-10.30
    Bus 841 from Torp to Göteborg Central Station 10.35-11.41
    Train 434 from Göteborg C to Stockholm C 12.24-15.26
    (There are other options, see

  • Contact

    Student office
    Study counsellor (advanced level)